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Module name  
Tree Structure and Function 
Forest Ecology and Management
Module coordinator 
 
Prof. Dr. Heinz Rennenberg    Email:  heinz.rennenberg@ctp.uni-freiburg.de 
Additional teaching staff 
Prof. Dr. Siegfried Fink, Dr. Jürgen Kreuzwieser 
Syllabus 
- Structure of roots, mechanisms of water and nutrient uptake by roots; significance of 
mycorrhizal symbiosis 
- Structure of xylem and phloem; xylem and phloem transport and transpiration  
- Leaf structure; CO2-exchange and photosynthesis in woody plants; tree respiration 
- Meristems and growth (cambium, shoot and root primordia, differentiation) 
- Source/sink relations in trees and its seasonality 
- Physiological basis of carbon fluxes in forest ecosystems   
- nutrient requirements of trees, consequences of nutrient deficiency and excess 
- regulation of tree nutrition 
- nutrient storage and mobilization  
Learning goals and qualifications  
The students will: 

achieve an in depth understanding of carbon relations of trees from the molecular via the 
physiological, eco-physiological and tree to the stand level. 

learn the role of trees in water relations of forest and the mechanisms involved in water 
acquisition, water transport inside the tree and water vapour flux into the atmosphere.  

obtain a detailed understanding of nutrient requirements of trees, nutrient aquisition, the 
mechanisms involved and its regulation 

understand the relations between structural aspects at the cell, tissue and organ level 
and the respective physiological functions  

become competent in linking growth processes at the cell and tissue level to “classical” 
growth parameters used in forestry (annual rings, volume yield etc.) 
  

 
Teaching and learning methods 
Lectures, tutorials  
Prerequisites 
none 
Requirements for registration 
none 
Distribution of work load  
Contact hours               80 h (Lectures, pracs, excursion, exam) 
Student learning          45 h (Preparation, reading etc.) 
Proposed assessment 
 
Link to learning resources   
 
Preliminary Reading 
Marschner H (1995) Mineral Nutrition of Higher Plants. Academic Press, London 
Landsberg JJ and Gower ST (1997) Applications of Physiological Ecology to Forest 
Management. Academic Press, San Diego 
Dickison, W.C. (2000): Integrative Plant Anatomy. Academic Press, San Diego 
Tyree, M.T. and M.H. Zimmermann (2002): Xylem Structure and the Ascent of Sap. 2nd. Ed. 
Springer, Berlin 
Comments 
 
 

Basic Information

Course:

Forest Ecology and Management

Degree:

Master of Science (M.Sc.)

Language of Instruction:

English

Duration:

2 years / 4 semesters

Total number of credits:

120 ECTS

accredited by: Acquin-Logo

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